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<p style="direction: ltr; text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">The University of Pennsylvania is home to a dynamic, interdisciplinary community of intellectually
adventurous researchers studying the brain using computational, theoretical, and experimental approaches. The Computational Neuroscience Initiative (CNI) offers a Graduate Certificate in Computational Neuroscience for PhD students in Neuroscience, Bioengineering,
Psychology, and Physics. The certificate provides training in theoretical models, neuronal and behavioral data analysis, predictive modeling, machine learning, and related computational methods. Trainees participate in coursework, lab rotations, seminars,
journal clubs, workshops, and a research-based capstone experience.</span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Learn more about CNI:
<a href="https://cni.upenn.edu/education" data-rich-text-autolink="" data-outlook-id="d5260942-30e3-429f-ac88-a1ee655ffcee" style="margin-top: 0px; margin-bottom: 0px;">
https://cni.upenn.edu/education</a></span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">You can find a (non-exhaustive) list of CNI-affiliated faculty here:
<a href="https://cni.upenn.edu/people" data-outlook-id="fda11a66-622f-414b-87d5-6131836080b0" style="margin-top: 0px; margin-bottom: 0px;">
https://cni.upenn.edu/people</a> . Additionally, Penn Mindcore brings together researchers across many disciplines for the integrative study of the mind:
<a href="https://mindcore.sas.upenn.edu/people/mindcore-faculty-and-associates/" data-outlook-id="85b012ca-4ca9-4450-807c-00e6f3ab6ac8" style="margin-top: 0px; margin-bottom: 0px;">
https://mindcore.sas.upenn.edu/people/mindcore-faculty-and-associates/</a></span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Additionally, if you’re interested specifically in computation in the audition and communication, the Computational
Approaches to the Neuroscience of Audition and Communication (CANAC) program is an NIH-funded training program for students interested in the neuroscience of hearing, audition, speech, and communication. CANAC combines experimental and theoretical training
through a collaborative, cross-disciplinary environment. Trainees are co-mentored by computational and experimental faculty, with opportunities for clinical and translational perspectives. Accepted trainees receive two years of funding for PhD thesis research,
as well as individualized advising on training and career development.</span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Learn more about CANAC:
<a href="https://www.med.upenn.edu/canac/" data-rich-text-autolink="" data-outlook-id="94dd2558-00e8-42bc-a111-994434659788" style="margin-top: 0px; margin-bottom: 0px;">
https://www.med.upenn.edu/canac/</a></span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Students interested in these programs or more generally in pursuing PhD research in computational and systems
neuroscience must first apply to the graduate program that best matches their interests:</span></p>
<ul data-spread="true" style="text-align: left;">
<li style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">
<p role="presentation">Neuroscience Graduate Group (NGG), Biomedical Graduate Studies (BGS)<br>
<a href="https://www.med.upenn.edu/ngg/admissions/" data-outlook-id="60f06250-7fed-4801-b19a-673a78ce2fb3" style="margin-top: 0px; margin-bottom: 0px;">Program information and application</a><br>
Deadline: December 1, 2026</p>
</li><li style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">
<p role="presentation">Bioengineering, School of Engineering and Applied Science (SEAS)<br>
<a href="https://be.engineering.upenn.edu/academics/doctoral/" data-outlook-id="b1d23f53-2322-4a86-be49-cf8c4ced9c2b" style="margin-top: 0px; margin-bottom: 0px;">Program information</a> ·
<a href="https://gradadm.engineering.upenn.edu/" data-outlook-id="eca9f17b-7e73-4fbb-8dbd-134bbd02c1ec" style="margin-top: 0px; margin-bottom: 0px;">
SEAS application portal</a><br>
Deadline: December 15, 2026</p>
</li><li style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">
<p role="presentation">Psychology, School of Arts and Sciences (SAS)<br>
<a href="https://psychology.sas.upenn.edu/graduate/information-applicants" data-outlook-id="874b67cd-c9b7-4870-ba0d-39c1cbda9ae0" style="margin-top: 0px; margin-bottom: 0px;">Program information and application</a><br>
Deadline: December 8, 2026</p>
</li><li style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">
<p role="presentation">Physics, School of Arts and Sciences (SAS)<br>
<a href="https://www.physics.upenn.edu/graduate/" data-outlook-id="90bdf961-95fc-4398-bd16-9faa0b7ab0aa" style="margin-top: 0px; margin-bottom: 0px;">Graduate program information</a> ·
<a href="https://www.physics.upenn.edu/graduate/admissions/" data-outlook-id="db44e1c3-1b67-480c-81d4-61793df82609" style="margin-top: 0px; margin-bottom: 0px;">
Physics graduate admissions</a><br>
Deadline: December 8, 2026</p>
</li></ul>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Admissions are handled separately by each graduate program, so applicants should apply directly to the program
that best fits their background and goals. Penn’s graduate programs are highly interdisciplinary, and students can pursue research across departmental and school boundaries.</span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Applicants should identify relevant Penn faculty and programs in their application and describe their interests
in computational, theoretical, experimental, auditory, systems, or cognitive neuroscience. We encourage students from diverse quantitative and scientific backgrounds to consider Penn for graduate study.</span></p>
<p style="text-align: left; text-indent: 0px;"><span style="font-family: Arial, Arial, Helvetica, sans-serif; font-size: 12pt; color: rgb(0, 0, 0);">Please check each program’s admissions page for any updates to deadlines, requirements, and application procedures.</span></p>
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<span style="text-transform: none;">________________________________</span></div>
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Maria Neimark Geffen, Ph.D. (she/her)</div>
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Professor</div>
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Co-director, Penn Computational Neuroscience Initiative</div>
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Departments of Otorhinolaryngology: HNS (primary), Neuroscience and Neurology</div>
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Perelman School of Medicine</div>
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University of Pennsylvania</div>
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5 Ravdin</div>
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3400 Spruce St.</div>
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Philadelphia PA 19104</div>
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Tel.: 215.898.0782 </div>
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<a href="https://pennmedicine.zoom.us/my/mgeffen?pwd=eWVFbUJFUGRJZ3g1eU9YZEZic2VwZz09" title="https://pennmedicine.zoom.us/my/mgeffen?pwd=eWVFbUJFUGRJZ3g1eU9YZEZic2VwZz09" data-outlook-id="1a8e1517-0223-461c-95ce-091d8e25ce1b" style="color: rgb(113, 79, 188);">My
zoom</a></div>
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<a href="https://calendar.app.google/upBGPWmGeHX5HtPc7" data-outlook-id="2afc439d-f593-4ec3-b8fe-0a7c39fb2e2f">My booking page</a></div>
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<a href="mailto:mgeffen@pennmedicine.upenn.edu" title="mailto:mgeffen@pennmedicine.upenn.edu" data-outlook-id="2c65bd20-702c-4456-ae94-cc77799a82e3" style="color: rgb(113, 79, 188);">mgeffen@pennmedicine.upenn.edu</a></div>
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<a href="http://hosting.med.upenn.edu/hearing" title="http://hosting.med.upenn.edu/hearing" data-outlook-id="c8323949-27d3-485a-b87e-89082a5f8339" style="color: rgb(113, 79, 188);">http://hosting.med.upenn.edu/hearing</a></div>
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<span style="text-transform: none;"><a href="https://bsky.app/profile/geffenlab.bsky.social" title="https://bsky.app/profile/geffenlab.bsky.social" data-outlook-id="5755dbac-f0a8-473d-8747-e1549bc8fa55" style="color: blue;">https://bsky.app/profile/geffenlab.bsky.social</a></span></div>
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<a href="https://www.linkedin.com/in/maria-geffen-bb6098380/" data-outlook-id="baaeb8eb-729f-43c2-9229-f4a5bde1172d">https://www.linkedin.com/in/maria-geffen-bb6098380/</a></div>
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